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Cross-modal reorganization of horizontal connectivity in auditory cortex without altering thalamocortical projections
Journal article   Open access   Peer reviewed

Cross-modal reorganization of horizontal connectivity in auditory cortex without altering thalamocortical projections

W J Gao and S L Pallas
The Journal of neuroscience, v 19(18), pp 7940-7950
15 Sep 1999
PMID: 10479695
url
https://doi.org/10.1523/JNEUROSCI.19-18-07940.1999View
Published, Version of Record (VoR) Open

Abstract

Animals, Newborn Afferent Pathways - growth & development Visual Pathways - growth & development Auditory Cortex - growth & development Axons - physiology Auditory Cortex - physiology Auditory Cortex - anatomy & histology Ferrets Retina - physiology Visual Pathways - physiology Thalamus - growth & development Animals Thalamus - physiology Afferent Pathways - physiology Aging Cerebral Cortex - growth & development Cerebral Cortex - physiology
The development of the different, highly specialized regions of the mammalian cerebral cortex depends in part on neural activity, either intrinsic spontaneous activity or externally driven sensory activity. To determine whether patterned sensory activity instructs the development of intrinsic cortical circuitry, we have experimentally altered the modality of sensory inputs to cerebral cortex. Neonatal diversion of retinal axons to the auditory thalamus (cross-modal rewiring) results in a primary auditory cortex (AI) that resembles visual cortex in its response properties and topography (Roe et al., 1990, 1992). To test the hypothesis that the visual response properties are created by a visually driven reorganization of auditory cortical circuitry, we investigated the effect of early visual experience on the development of intrinsic, horizontal connections within AI. Horizontal connections are likely to play an important role in the construction of visual response properties in AI as they do in visual cortex. Here we show that early visual inputs to auditory thalamus can reorganize horizontal connections in AI, causing both an increase in their extent and a change in pattern, so that projections are not restricted to the isofrequency axis, but extend in a more isotropic pattern around the injection site. Thus, changing afferent modality, without altering the source of the thalamocortical axons, can profoundly alter cortical circuitry. Similar changes may underlie cortical compensatory processes in deaf or blind humans and may also have played a role in the parcellation of neocortex during mammalian evolution.

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