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Elevated Levels of the Plasmodium yoelii Homologue of Macrophage Migration Inhibitory Factor Attenuate Blood-Stage Malaria
Journal article   Open access   Peer reviewed

Elevated Levels of the Plasmodium yoelii Homologue of Macrophage Migration Inhibitory Factor Attenuate Blood-Stage Malaria

Swati Thorat, Thomas M Daly, Lawrence W Bergman and James M Burns
Infection and immunity, v 78(12), pp 5151-5162
Dec 2010
PMID: 20837716
url
https://doi.org/10.1128/IAI.00277-10View
Published, Version of Record (VoR) Open

Abstract

Microbial Immunity and Vaccines
The excessive production of proinflammatory cytokines plays a significant role in the pathogenesis of severe malaria. Mammalian macrophage migration inhibitory factor (MIF) (mMIF) is an immune mediator that promotes a sustained proinflammatory response by inhibiting the glucocorticoid-mediated downregulation of inflammation. In addition, Plasmodium parasites also encode a homologue of mammalian MIF that is expressed in asexual-stage parasites. We used the Plasmodium yoelii murine model to study the potential role of parasite-encoded MIF in the pathogenesis of malaria. Antibodies raised against purified, non-epitope-tagged P. yoelii MIF ( Py MIF) were used to localize expression in trophozoite- and schizont-stage parasites and demonstrate extracellular release. In vitro , recombinant Py MIF was shown to actively induce the chemotaxis of macrophages but did not induce or enhance tumor necrosis factor alpha (TNF-α) production from peritoneal macrophages. To examine the role of parasite-derived Py MIF in vivo , two transgenic parasite lines that constitutively overexpress Py MIF were generated, one in a nonlethal P. yoelii 17X background [ Py 17X-MIF(+)] and the other in a lethal P. yoelii 17XL background [ Py 17XL-MIF(+)]. Challenge studies with transgenic parasites in mice showed that the increased expression of Py MIF resulted in a reduction in disease severity. Mice infected with Py 17X-MIF(+) developed lower peak parasitemia levels than controls, while malaria-associated anemia was unaltered. Infection with Py 17XL-MIF(+) resulted in a prolonged course of infection and a reduction in the overall mortality rate. Combined, the data indicate that parasite-derived MIF does not contribute significantly to immunopathology but, through its chemotactic ability toward macrophages, may attenuate disease and prolong infection of highly virulent parasite isolates.

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Web of Science research areas
Immunology
Infectious Diseases
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