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Locus Control Region Transcription Plays an Active Role in Long-Range Gene Activation
Journal article   Open access   Peer reviewed

Locus Control Region Transcription Plays an Active Role in Long-Range Gene Activation

Yugong Ho, Felice Elefant, Stephen A Liebhaber and Nancy E Cooke
Molecular cell, v 23(3), pp 365-375
2006
PMID: 16885026
url
https://doi.org/10.1016/j.molcel.2006.05.041View
Published, Version of Record (VoR) Open

Abstract

DNA
Activation of eukaryotic genes often relies on remote chromatin determinants. How these determinants function remains poorly understood. The hGH gene is activated by a 5′-remote locus control region (LCR). Pituitary-specific DNase I hypersensitive site I (HSI), the dominant hGH LCR element, is separated from the hGH-N promoter by a 14.5 kb span that encompasses the B-lymphocyte-specific CD79b gene. Here, we describe a domain of noncoding Pol II transcription in pituitary somatotropes that includes the hGH LCR and adjacent CD79b locus. This entire “LCR domain of transcription” is HIS dependent and terminates 3′ to CD79b, leaving a gap in transcription between this domain and the target hGH-N promoter. Insertion of a Pol II terminator within the LCR blocks CD79b transcription and represses hGH-N expression. These data document an essential role for LCR transcription in long-range control, link “bystander” CD79b transcription to this process, and support a unique model for locus activation.

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Web of Science research areas
Biochemistry & Molecular Biology
Cell Biology
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