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MAP kinase subcellular localization controls both pattern and proliferation in the developing Drosophila wing
Journal article   Open access   Peer reviewed

MAP kinase subcellular localization controls both pattern and proliferation in the developing Drosophila wing

Daniel R Marenda, Alysia D Vrailas, Aloma B Rodrigues, Summer Cook, Maureen A Powers, James A Lorenzen, Lizabeth A Perkins and Kevin Moses
Development (Cambridge), v 133(1), pp 43-51
Jan 2006
PMID: 16308331
url
https://doi.org/10.1242/dev.02168View
Published, Version of Record (VoR) Open

Abstract

Immunohistochemistry Phosphorylation Cell Proliferation Drosophila Cytoplasm - metabolism Protein Transport - physiology Wings, Animal - enzymology Hot Temperature HSP70 Heat-Shock Proteins - metabolism Animals Cell Nucleus - metabolism Flow Cytometry Wings, Animal - growth & development Signal Transduction - physiology Cell Differentiation - physiology Mitogen-Activated Protein Kinases - metabolism
Mitogen-activated protein kinases (MAPKs) phosphorylate target proteins in both the cytoplasm and nucleus, and a strong correlation exists between the subcellular localization of MAPK and resulting cellular responses. It was thought that MAPK phosphorylation was always followed by rapid nuclear translocation. However, we and others have found that MAPK phosphorylation is not always sufficient for nuclear translocation in vivo. In the developing Drosophila wing, MAPK-mediated signaling is required both for patterning and for cell proliferation, although the mechanism of this differential control is not fully understood. Here, we show that phosphorylated MAPK (pMAPK) is held in the cytoplasm in differentiating larval and pupal wing vein cells, and we show that this cytoplasmic hold is required for vein cell fate. At the same time, we show that MAPK does move into the nucleus of other wing cells where it promotes cell proliferation. We propose a novel Ras pathway bifurcation in Drosophila and our results suggest a mechanism by which MAPK phosphorylation can signal two different cellular outcomes (differentiation versus proliferation) based on the subcellular localization of MAPK.

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Collaboration types
Domestic collaboration
Web of Science research areas
Developmental Biology
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