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The importance of vibronic perturbations in ferrocytochrome c spectra: A reevaluation of spectral properties based on low-temperature optical absorption, resonance Raman, and molecular-dynamics simulations
Journal article   Peer reviewed

The importance of vibronic perturbations in ferrocytochrome c spectra: A reevaluation of spectral properties based on low-temperature optical absorption, resonance Raman, and molecular-dynamics simulations

Matteo Levantino, Qing Huang, Antonio Cupane, Monique Laberge, Andrew Hagarman and Reinhard Schweitzer-Stenner
The Journal of chemical physics, v 123(5), pp 054508-054508-12
09 Aug 2005
PMID: 16108670

Abstract

We have measured and analyzed the low-temperature ( T = 10 K ) absorption spectrum of reduced horse heart and yeast cytochrome c. Both spectra show split and asymmetric Q 0 and Q v bands. The spectra were first decomposed into the individual split vibronic sidebands assignable to B 1 g ( ν 15 ) and A 2 g ( ν 19 , ν 21 , and ν 22 ) Herzberg-Teller active modes due to their strong intensity in resonance Raman spectra acquired with Q 0 and Q v excitations. The measured band splittings and asymmetries cannot be rationalized solely in terms of electronic perturbations of the heme macrocycle. On the contrary, they clearly point to the importance of considering not only electronic perturbations but vibronic perturbations as well. The former are most likely due to the heterogeneity of the electric field produced by charged side chains in the protein environment, whereas the latter reflect a perturbation potential due to multiple heme-protein interactions, which deform the heme structure in the ground and excited states. Additional information about vibronic perturbations and the associated ground-state deformations are inferred from the depolarization ratios of resonance Raman bands. The results of our analysis indicate that the heme group in yeast cytochrome c is more nonplanar and more distorted along a B 2 g coordinate than in horse heart cytochrome c. This conclusion is supported by normal structural decomposition calculations performed on the heme extracted from molecular-dynamic simulations of the two investigated proteins. Interestingly, the latter are somewhat different from the respective deformations obtained from the x-ray structures.

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Collaboration types
Domestic collaboration
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Web of Science research areas
Chemistry, Physical
Physics, Atomic, Molecular & Chemical
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