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p56Lck Tyrosine Kinase Enhances the Assembly of Death-inducing Signaling Complex during Fas-mediated Apoptosis
Journal article   Open access   Peer reviewed

p56Lck Tyrosine Kinase Enhances the Assembly of Death-inducing Signaling Complex during Fas-mediated Apoptosis

Ehssan Sharif-Askari, Denis Gaucher, Rabih Halwani, Jennifer Ma, Kevin Jao, Ali Abdallah, Elias K. Haddad and Rafick-Pierre Sékaly
The Journal of biological chemistry, v 282(49), pp 36048-36056
07 Dec 2007
PMID: 17932036
url
http://www.jbc.org/content/282/49/36048.full.pdfView
Published, Version of Record (VoR) Open
url
https://doi.org/10.1074/jbc.M706007200View
Published, Version of Record (VoR) Open

Abstract

Although the death-inducing signaling complex (DISC) is rapidly assembled, several lines of evidence suggest that formation of this complex is not the first consequence of cell surface CD95 (Fas) stimulation but rather a later step in this process. Activation of Fas triggers a cascade of signaling events that culminate in cellular apoptosis. Tyrosine kinases are critical effectors in T cell activation. However, their functional involvement in death receptor-mediated apoptosis is unknown. Here, we used p56Lck-deficient cells to show that CD95-induced cell death is highly dependent on p56Lck activity and its localization within plasma membrane. We found that p56Lck acts upstream of the mitochondria; in the absence of p56Lck, Bid cleavage and the release of cytochrome c were severely impaired. Moreover, p56Lck-deficient cells or cells expressing an inactive form of p56Lck displayed defective formation of the DISC post CD95 stimulation. In vivo reconstitution of thymocytes from p56lck-deficient mice, which are resistant to apoptosis, with p56Lck restored Fas-mediated cell death. Our results support a novel model whereby sensitivity to apoptosis is regulated through quantitative changes in the stoichiometry of DISC components triggered by p56Lck activation and localization.

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Collaboration types
Domestic collaboration
Web of Science research areas
Biochemistry & Molecular Biology
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